Within the fossil sample, configurations of key sulci delineate distinct morphotypes spanning from more ancestral to more derived arrangements, highlighting anatomical diversity within assemblages that include taxa under debate.

Considered alongside the comparative variability patterns, these results support a mosaic model of evolution, in which cortical reorganisation unfolded in stages and differed across lineages rather than following a single linear trajectory. This research demonstrates how advanced digital methods, anchored by a rigorously characterised comparative framework, refine inferences about cerebral change through time.